Part 8 (2/2)

H_{2}SO_{4} destruction was almost complete. In these cases there was no question of a mere oxidation to oxyluciferin, as no oxyluciferin could be demonstrated after boiling with such strong acids. An actual destruction, probably an hydrolysis of the luciferin molecule, occurred.

We shall have occasion to refer to this again in considering the protein nature of luciferin. It must be borne in mind that many proteins require four or five days' boiling with 20 per cent. HCl for complete hydrolysis to amino-acids. Luciferin forms a solution in water, probably colloidal, although the luciferin will dialyze through parchment or collodion membranes. It is rather readily adsorbed by various finely divided materials such as bone black, Fe(OH)_{3}, kaolin, talc and CaCo_{3}. It is not destroyed by any of the enzyme solutions which I have tried.

These include such as are widely divergent in action: pepsin HCl, trypsin, erepsin, salivary and malt diastase, yeast invertase, urease, rennin and the enzymes of dried spleen, kidney and liver substances.

By extracting the dried Cypridinas ground to a powder, the solubility of luciferin in non-aqueous solvents could be easily studied, and by adding such reagents as dilute acids, alkalies, neutral salts and the alkaloidal reagents to an aqueous solution of luciferin the general biochemical behavior of luciferin can be quite accurately stated. For convenience the results of this study are given in Table 8.

TABLE 8

_Properties of Photogenic Substances from Cypridina_

========================================================================= Property

Luciferase

Luciferin ------------------------+-----------------------+------------------------ _Salting out_

By saturation NaCl

Not precipitated

Not precipitated.

By half saturation

Do.

Do.

MgSO_{4}

By saturation MgSO_{4}

Nearly completely

Partially

precipitated

precipitated.

By saturation MgSO_{4}

...

Do.

+ acetic acid

By half saturation

(NH_{4})_{2}SO_{4}

Slightly precipitated

Not precipitated.

By saturation

Completely precipitated

Nearly completely (NH_{4})_{2}SO_{4}

precipitated.

By saturation

(NH_{4})_{2}SO_{4} +

...

Nearly completely acetic acid

precipitated.

_Solubility in_

Methyl alcohol

Insoluble

Soluble.

Ethyl alcohol

Do.

Do.

90 per cent.

Do.

Do.

70 per cent.

Do.

Do.

50 per cent.

Slightly soluble

Do.

Propyl alcohol

Insoluble

Do.

Isobutyl alcohol

Do.

Fairly soluble.

Amyl alcohol

Do.

Slightly soluble.

Benzyl alcohol

Do.

Soluble.

Acetone

Do.

Fairly soluble.

90 per cent.

Do.

Soluble.

70 per cent.

Slightly soluble

Do.

50 per cent.

Fairly soluble

Do.

Ethyl acetate

Insoluble

Do.

Ethyl propionate

Do.

Fairly soluble.

Ethyl butyrate

Do.

Do.

Ethyl valerate

Do.

Slightly soluble.

Ethyl nitrate

Do.

Very slightly soluble.

Glycerine

Do.

Soluble.

Glycol

Do.

Do.

Ether

Do.

Insoluble.

Chloroform

Do.

Do.

Carbon disulfide

Do.

Do.

Carbon tetrachloride

Do.

Do.

Benzol

Do.

Do.

Toluol

Do.

Do.

Xylol

Do.

Do.

Petroleum ether

Do.

Do.

Anilin

Do.

Do.

Glacial acetic acid

Do.

Fairly soluble.

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