Volume Ii Part 1 (1/2)

More Letters of Charles Darwin.

Volume II.

by Charles Darwin.

CHAPTER 2.VII.--GEOGRAPHICAL DISTRIBUTION.

1843-1882 (Continued) (1867-1882.)

LETTER 378. J.D. HOOKER TO CHARLES DARWIN. Kew, January 20th, 1867.

Prof. Miquel, of Utrecht, begs me to ask you for your carte, and offers his in return. I grieve to bother you on such a subject. I am sick and tired of this carte correspondence. I cannot conceive what Humboldt's Pyrenean violet is: no such is mentioned in Webb, and no alpine one at all. I am sorry I forgot to mention the stronger African affinity of the eastern Canary Islands. Thank you for mentioning it. I cannot admit, without further a.n.a.lysis, that most of the peculiar Atlantic Islands genera were derived from Europe, and have since become extinct there.

I have rather thought that many are only altered forms of existing European genera; but this is a very difficult point, and would require a careful study of such genera and allies with this object in view. The subject has often presented itself to me as a grand one for a.n.a.lytic botany. No doubt its establishment would account for the community of the peculiar genera on the several groups and islets, but whilst so many species are common we must allow for a good deal of migration of peculiar genera too.

By Jove! I will write out next mail to the Governor of St. Helena for boxes of earth, and you shall have them to grow. Thanks for telling me of having suggested to me the working out of proportions of plants with irregular flowers in islands. I thought it was a deuced deal too good an idea to have arisen spontaneously in my block, though I did not recollect your having done so. No doubt your suggestion was crystallised in some corner of my sensorium. I should like to work out the point.

Have you Kerguelen Land amongst your volcanic islands? I have a curious book of a sealer who was wrecked on the island, and who mentions a volcanic mountain and hot springs at the S.W. end; it is called the ”Wreck of the Favourite.” (378/1. ”Narrative of the Wreck of the 'Favourite' on the Island of Desolation; detailing the Adventures, Sufferings and Privations of John Munn; an Historical Account of the Island and its Whale and Sea Fisheries.” Edited by W.B. Clarke: London, 1850.)

LETTER 379. TO J.D. HOOKER. Down, March 17th, 1867.

It is a long time since I have written, but I cannot boast that I have refrained from charity towards you, but from having lots of work...You ask what I have been doing. Nothing but blackening proofs with corrections. I do not believe any man in England naturally writes so vile a style as I do...

In your paper on ”Insular Floras” (page 9) there is what I must think an error, which I before pointed out to you: viz., you say that the plants which are wholly distinct from those of nearest continent are often very common instead of very rare. (379/1. ”Insular Floras,” pamphlet reprinted from the ”Gardeners' Chronicle,” page 9: ”As a general rule the species of the mother continent are proportionally the most abundant, and cover the greatest surface of the islands. The peculiar species are rarer, the peculiar genera of continental affinity are rarer still; whilst the plants having no affinity with those of the mother continent are often very common.” In a letter of March 20th, 1867, Sir Joseph explains that in the case of the Atlantic islands it is the ”peculiar genera of EUROPEAN AFFINITY that are so rare,” while Clethra, Dracaena and the Laurels, which have no European affinity, are common.) Etty (379/2. Mr. Darwin's daughter, now Mrs. Litchfield.), who has read your paper with great interest, was confounded by this sentence. By the way, I have stumbled on two old notes: one, that twenty-two species of European birds occasionally arrive as chance wanderers to the Azores; and, secondly, that trunks of American trees have been known to be washed on the sh.o.r.es of the Canary Islands by the Gulf-stream, which returns southward from the Azores. What poor papers those of A. Murray are in ”Gardeners' Chronicle.” What conclusions he draws from a single Carabus (379/3. ”Dr. Hooker on Insular Floras” (”Gardeners' Chronicle,”

1867, pages 152, 181). The reference to the Carabidous beetle (Aplothorax) is at page 181.), and that a widely ranging genus! He seems to me conceited; you and I are fair game geologically, but he refers to Lyell, as if his opinion on a geological point was worth no more than his own. I have just bought, but not read a sentence of, Murray's big book (379/4. ”Geographical Distribution of Mammals,” 1866.), second-hand, for 30s., new, so I do not envy the publishers. It is clear to me that the man cannot reason. I have had a very nice letter from Scott at Calcutta (379/5. See Letter 150.): he has been making some good observations on the acclimatisation of seeds from plants of same species, grown in different countries, and likewise on how far European plants will stand the climate of Calcutta. He says he is astonished how well some flourish, and he maintains, if the land were unoccupied, several could easily cross, spreading by seed, the Tropics from north to south, so he knows how to please me; but I have told him to be cautious, else he will have dragons down on him...

As the Azores are only about two-and-a-half times more distant from America (in the same lat.i.tude) than from Europe, on the occasional migration view (especially as oceanic currents come directly from West Indies and Florida, and heavy gales of wind blow from the same direction), a large percentage of the flora ought to be American; as it is, we have only the Sanicula, and at present we have no explanation of this apparent anomaly, or only a feeble indication of an explanation in the birds of the Azores being all European.

LETTER 380. TO J.D. HOOKER. Down, March 21st [1867].

Many thanks for your pleasant and very amusing letter. You have been treated shamefully by Etty and me, but now that I know the facts, the sentence seems to me quite clear. Nevertheless, as we have both blundered, it would be well to modify the sentence something as follows: ”whilst, on the other hand, the plants which are related to those of distant continents, but have no affinity with those of the mother continent, are often very common.” I forget whether you explain this circ.u.mstance, but it seems to me very mysterious (380/1. Sir Joseph Hooker wrote (March 23rd, 1867): ”I see you 'smell a rat' in the matter of insular plants that are related to those of [a] distant continent being common. Yes, my beloved friend, let me make a clean breast of it. I only found it out after the lecture was in print!...I have been waiting ever since to 'think it out,' and write to you about it, coherently. I thought it best to squeeze it in, anyhow or anywhere, rather than leave so curious a fact unnoticed.”)...Do always remember that nothing in the world gives us so much pleasure as seeing you here whenever you can come. I chuckle over what you say of And. Murray, but I must grapple with his book some day.

LETTER 381. TO C. LYELL. Down, October 31st [1867].

Mr. [J.P. Mansel] Weale sent to me from Natal a small packet of dry locust dung, under 1/2 oz., with the statement that it is believed that they introduce new plants into a district. (381/1. See Volume I., Letter 221.) This statement, however, must be very doubtful. From this packet seven plants have germinated, belonging to at least two kinds of gra.s.ses. There is no error, for I dissected some of the seeds out of the middle of the pellets. It deserves notice that locusts are sometimes blown far out to sea. I caught one 370 miles from Africa, and I have heard of much greater distances. You might like to hear the following case, as it relates to a migratory bird belonging to the most wandering of all orders--viz. the woodc.o.c.k. (381/2. ”Origin,” Edition VI., page 328.) The tarsus was firmly coated with mud, weighing when dry 9 grains, and from this the Juncus bufonius, or toad rush, germinated. By the way, the locust case verifies what I said in the ”Origin,” that many possible means of distribution would be hereafter discovered. I quite agree about the extreme difficulty of the distribution of land mollusca. You will have seen in the last edition of ”Origin” (381/3. ”Origin,” Edition IV., page 429. The reference is to MM. Marten's (381/4. For Marten's read Martins' [the name is wrongly spelt in the ”Origin of Species.”]) experiments on seeds ”in a box in the actual sea.”) that my observations on the effects of sea-water have been confirmed. I still suspect that the legs of birds which roost on the ground may be an efficient means; but I was interrupted when going to make trials on this subject, and have never resumed it.

We shall be in London in the middle of latter part of November, when I shall much enjoy seeing you. Emma sends her love, and many thanks for Lady Lyell's note.

LETTER 382. TO J.D. HOOKER. Down, Wednesday [1867].

I daresay there is a great deal of truth in your remarks on the glacial affair, but we are in a muddle, and shall never agree. I am bigoted to the last inch, and will not yield. I cannot think how you can attach so much weight to the physicists, seeing how Hopkins, Hennessey, Haughton, and Thomson have enormously disagreed about the rate of cooling of the crust; remembering Herschel's speculations about cold s.p.a.ce (382/1.

The reader will find some account of Herschel's views in Lyell's ”Principles,” 1872, Edition XI., Volume I., page 283.), and bearing in mind all the recent speculations on change of axis, I will maintain to the death that your case of Fernando Po and Abyssinia is worth ten times more than the belief of a dozen physicists. (382/2. See ”Origin,”

Edition VI., page 337: ”Dr. Hooker has also lately shown that several of the plants living on the upper parts of the lofty island of Fernando Po and on the neighbouring Cameroon mountains, in the Gulf of Guinea, are closely related to those in the mountains of Abyssinia, and likewise to those of temperate Europe.” Darwin evidently means that such facts as these are better evidence of the gigantic periods of time occupied by evolutionary changes than the discordant conclusions of the physicists.

See ”Linn. Soc. Journ.” Volume VII., page 180, for Hooker's general conclusions; also Hooker and Ball's ”Marocco,” Appendix F, page 421. For the case of Fernando Po see Hooker (”Linn. Soc. Journ.” VI., 1861, page 3, where he sums up: ”Hence the result of comparing Clarence Peak flora [Fernando Po] with that of the African continent is--(1) the intimate relations.h.i.+p with Abyssinia, of whose flora it is a member, and from which it is separated by 1800 miles of absolutely unexplored country; (2) the curious relations.h.i.+p with the East African islands, which are still farther off; (3) the almost total dissimilarity from the Cape flora.” For Sir J.D. Hooker's general conclusions on the Cameroon plants see ”Linn. Soc. Journ.” VII., page 180. More recently equally striking cases have come to light: for instance, the existence of a Mediterranean genus, Adenocarpus, in the Cameroons and on Kilima Njaro, and nowhere else in Africa; and the probable migration of South African forms along the highlands from the Natal District to Abysinnia. See Hooker, ”Linn.

Soc. Journ.” XIV., 1874, pages 144-5.) Your remarks on my regarding temperate plants and disregarding the tropical plants made me at first uncomfortable, but I soon recovered. You say that all botanists would agree that many tropical plants could not withstand a somewhat cooler climate. But I have come not to care at all for general beliefs without the special facts. I have suffered too often from this: thus I found in every book the general statement that a host of flowers were fertilised in the bud, that seeds could not withstand salt water, etc., etc. I would far more trust such graphic accounts as that by you of the mixed vegetation on the Himalayas and other such accounts. And with respect to tropical plants withstanding the slowly coming on cool period, I trust to such facts as yours (and others) about seeds of the same species from mountains and plains having acquired a slightly different climatal const.i.tution. I know all that I have said will excite in you savage contempt towards me. Do not answer this rigmarole, but attack me to your heart's content, and to that of mine, whenever you can come here, and may it be soon.

LETTER 383. J.D. HOOKER TO CHARLES DARWIN. Kew, 1870.

(383/1. The following extract from a letter of Sir J.D. Hooker shows the tables reversed between the correspondents.)