Volume I Part 25 (2/2)
LETTER 142. H. FALCONER TO CHARLES DARWIN.
(142/1. This refers to the MS. of Falconer's paper ”On the American Fossil Elephant of the Regions bordering the Gulf of Mexico (E. Columbi, Falc.),” published in the ”Natural History Review,” January, 1863, page 43. The section dealing with the bearing of his facts on Darwin's views is at page 77. He insists strongly (page 78) on the ”persistence and uniformity of the characters of the molar teeth in the earliest known mammoth, and his most modern successor.” Nevertheless, he adds that the ”inferences I draw from these facts are not opposed to one of the leading propositions of Darwin's theory.” These admissions were the more satisfactory since, as Falconer points out (page 77), ”I have been included by him in the category of those who have vehemently maintained the persistence of specific characters.”)
21, Park Crescent, Portland Place, N.W., September 24th [1862].
Do not be frightened at the enclosure. I wish to set myself right by you before I go to press. I am bringing out a heavy memoir on elephants--an omnium gatherum affair, with observations on the fossil and recent species. One section is devoted to the persistence in time of the specific characters of the mammoth. I trace him from before the Glacial period, through it and after it, unchangeable and unchanged as far as the organs of digestion (teeth) and locomotion are concerned. Now, the Glacial period was no joke: it would have made ducks and drakes of your dear pigeons and doves.
With all my shortcomings, I have such a sincere and affectionate regard for you and such admiration of your work, that I should be pained to find that I had expressed my honest convictions in a way that would be open to any objection by you. The reasoning may be very stupid, but I believe that the observation is sound. Will you, therefore, look over the few pages which I have sent, and tell me whether you find any flaw, or whether you think I should change the form of expression? You have been so unhandsomely and uncandidly dealt with by a friend of yours and mine that I should be sorry to find myself in the position of an opponent to you, and more particularly with the chance of making a fool of myself.
I met your brother yesterday, who tells me you are coming to town. I hope you will give me a hail. I long for a jaw with you, and have much to speak to you about.
You will have seen the eclairciss.e.m.e.nt about the Eocene monkeys of England. By a touch of the conjuring wand they have been metamorphosed--a la Darwin--into Hyracotherian pigs. (142/2. ”On the Hyracotherian Character of the Lower Molars of the supposed Macacus from the Eocene Sand of Kyson, Suffolk.” ”Ann. Mag. Nat. Hist.” Volume X., 1862, page 240. In this note Owen stated that the teeth which he had named Macacus (”Ann. Mag.” 1840, page 191) most probably belonged to Hyracotherium cuniculus. See ”A Catalogue of British Fossil Vertebrata,”
A.S. Woodward and C.D. Sherborn, 1890, under Hyracotherium, page 356; also Zittel's ”Handbuch der Palaeontologie” Abth. I., Bd. IV., Leipzig, 1891-93, page 703.) Would you believe it? This even is a gross blunder.
They are not pigs.
LETTER 143. TO HUGH FALCONER. Down, October 1st [1862].
On my return home yesterday I found your letter and MS., which I have read with extreme interest. Your note and every word in your paper are expressed with the same kind feeling which I have experienced from you ever since I have had the happiness of knowing you. I value scientific praise, but I value incomparably higher such kind feeling as yours.
There is not a single word in your paper to which I could possibly object: I should be mad to do so; its only fault is perhaps its too great kindness. Your case seems the most striking one which I have met with of the persistence of specific characters. It is very much the more striking as it relates to the molar teeth, which differ so much in the species of the genus, and in which consequently I should have expected variation. As I read on I felt not a little dumbfounded, and thought to myself that whenever I came to this subject I should have to be savage against myself; and I wondered how savage you would be. I trembled a little. My only hope was that something could be made out of the bog N. American forms, which you rank as a geographical race; and possibly hereafter out of the Sicilian species. Guess, then, my satisfaction when I found that you yourself made a loophole (143/1. This perhaps refers to a pa.s.sage (”N.H. Review,” 1863, page 79) in which Falconer allows the existence of intermediate forms along certain possible lines of descent.
Falconer's reference to the Sicilian elephants is in a note on page 78; the bog-elephant is mentioned on page 79.), which I never, of course, could have guessed at; and imagine my still greater satisfaction at your expressing yourself as an unbeliever in the eternal immutability of species. Your final remarks on my work are too generous, but have given me not a little pleasure. As for criticisms, I have only small ones.
When you speak of ”moderate range of variation” I cannot but think that you ought to remind your readers (though I daresay previously done) what the amount is, including the case of the American bog-mammoth. You speak of these animals as having been exposed to a vast range of climatal changes from before to after the Glacial period. I should have thought, from a.n.a.logy of sea-sh.e.l.ls, that by migration (or local extinction when migration not possible) these animals might and would have kept under nearly the same climate.
A rather more important consideration, as it seems to me, is that the whole proboscidean group may, I presume, be looked at as verging towards extinction: anyhow, the extinction has been complete as far as Europe and America are concerned. Numerous considerations and facts have led me in the ”Origin” to conclude that it is the flouris.h.i.+ng or dominant members of each order which generally give rise to new races, sub-species, and species; and under this point of view I am not at all surprised at the constancy of your species. This leads me to remark that the sentence at the bottom of page [80] is not applicable to my views (143/2. See Falconer at the bottom of page 80: it is the old difficulty--how can variability co-exist with persistence of type? In our copy of the letter the pa.s.sage is given as occurring on page 60, a slip of the pen for page 80.), though quite applicable to those who attribute modification to the direct action of the conditions of life.
An elephant might be more individually variable than any known quadruped (from the effects of the conditions of life or other innate unknown causes), but if these variations did not aid the animal in better resisting all hostile influences, and therefore making it increase in numbers, there would be no tendency to the preservation and acc.u.mulation of such variations--i.e. to the formation of a new race. As the proboscidean group seems to be from utterly unknown causes a failing group in many parts of the world, I should not have antic.i.p.ated the formation of new races.
You make important remarks versus Natural Selection, and you will perhaps be surprised that I do to a large extent agree with you. I could show you many pa.s.sages, written as strongly as I could in the ”Origin,”
declaring that Natural Selection can do nothing without previous variability; and I have tried to put equally strongly that variability is governed by many laws, mostly quite unknown. My t.i.tle deceives people, and I wish I had made it rather different. Your phyllotaxis (143/3. Falconer, page 80: ”The law of Phyllotaxis...is nearly as constant in its manifestation as any of the physical laws connected with the material world.”) will serve as example, for I quite agree that the spiral arrangement of a certain number of whorls of leaves (however that may have primordially arisen, and whether quite as invariable as you state), governs the limits of variability, and therefore governs what Natural Selection can do. Let me explain how it arose that I laid so much stress on Natural Selection, and I still think justly. I came to think from geographical distribution, etc., etc., that species probably change; but for years I was stopped dead by my utter incapability of seeing how every part of each creature (a woodp.e.c.k.e.r or swallow, for instance) had become adapted to its conditions of life. This seemed to me, and does still seem, the problem to solve; and I think Natural Selection solves it, as artificial selection solves the adaptation of domestic races for man's use. But I suspect that you mean something further,--that there is some unknown law of evolution by which species necessarily change; and if this be so, I cannot agree. This, however, is too large a question even for so unreasonably long a letter as this. Nevertheless, just to explain by mere valueless conjectures how I imagine the teeth of your elephants change, I should look at the change as indirectly resulting from changes in the form of the jaws, or from the development of tusks, or in the case of the primigenius even from correlation with the woolly covering; in all cases Natural Selection checking the variation. If, indeed, an elephant would succeed better by feeding on some new kinds of food, then any variation of any kind in the teeth which favoured their grinding power would be preserved. Now, I can fancy you holding up your hands and crying out what bos.h.!.+ To return to your concluding sentence: far from being surprised, I look at it as absolutely certain that very much in the ”Origin” will be proved rubbish; but I expect and hope that the framework will stand. (143/4.
Falconer, page 80: ”He [Darwin] has laid the foundations of a great edifice: but he need not be surprised if, in the progress of erection, the superstructure is altered by his successors...”)
I had hoped to have called on you on Monday evening, but was quite knocked up. I saw Lyell yesterday morning. He was very curious about your views, and as I had to write to him this morning I could not help telling him a few words on your views. I suppose you are tired of the ”Origin,” and will never read it again; otherwise I should like you to have the third edition, and would gladly send it rather than you should look at the first or second edition. With cordial thanks for your generous kindness.
LETTER 144. J.D. HOOKER TO CHARLES DARWIN. Royal Gardens, Kew, November 7th, 1862.
I am greatly relieved by your letter this morning about my Arctic essay, for I had been conjuring up some egregious blunder (like the granitic plains of Patagonia).. Certes, after what you have told me of Dawson, he will not like the letter I wrote to him days ago, in which I told him that it was impossible to entertain a strong opinion against the Darwinian hypothesis without its giving rise to a mental twist when viewing matters in which that hypothesis was or might be involved. I told him I felt that this was so with me when I opposed you, and that all minds are subject to such obliquities!--the Lord help me, and this to an LL.D. and Princ.i.p.al of a College! I proceeded to discuss his Geology with the effrontery of a novice; and, thank G.o.d, I urged the very argument of your letter about evidence of subsidence--viz., not all submerged at once, and glacial action being subaerial and not oceanic.
Your letter hence was a relief, for I felt I was hardly strong enough to have launched out as I did to a professed geologist.
(144/1. [On the subject of the above letter, see one of earlier date by Sir J.D. Hooker (November 2nd, 1862) given in the present work (Letter 354) with Darwin's reply (Letter 355).])
LETTER 145. TO HUGH FALCONER. Down, November 14th [1862].
I have read your paper (145/1. ”On the disputed Affinity of the Mammalian Genus Plagiaulax, from the Purbeck beds.”--”Quart. Journ.
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