Part 9 (2/2)

To the same part of the choroid coat, where the ciliary ligament begins, is fixed a moveable and curious membrane, called the iris; this membrane has a perforation in the middle, called the pupil, for the admission of the rays of light. The iris is composed of two kinds of fibres: those of the one sort tend, like the radii of a circle, towards its centre, and the others form a number of concentric circles round the same centre. The pupil is of no constant magnitude, for when a very luminous object is viewed, the circular fibres of the iris contract, and diminish its orifice; and, on the contrary, when objects are dark and obscure, those fibres relax, and suffer the pupil to enlarge, in order to admit a greater quant.i.ty of light into the eye: it is thought that the radial fibres also a.s.sist in enlarging the pupil. The iris is variously coloured in different persons, but according to no certain rule; though in general, they who have light hair, and a fair complexion, have the iris blue or grey; and, on the contrary, they whose hair and complexion are dark, have the iris of a deep brown; but whether this difference in colour occasions any difference in the sense, is not yet discovered. In the human eye the whole choroid coat, and even the interior surface of the iris or uvea, is lined with a black mucus; this mucus, or as it is called, pigmentum, is darkest in young persons, and becomes more light coloured as we advance in years. In many animals, but more particularly those which catch their prey in the night, this pigmentum is of a bright colour: its use will appear afterwards.

The last, and innermost coat of the eye, is the retina, it differs much from the above mentioned coats, being very delicate and tender.

It is nothing but an expansion of the medullary part of the optic nerve, which is inserted into each eye, nearer the nose, and a little higher, than the axis. This coat has been thought by many to end where the choroides, going inwards, towards the axis of the eye, forms the ciliary ligament; Dr. Monro thinks that it is not continued so far, and we cannot see with what advantage it could have been continued to the ciliary ligament, since none of the rays of light, pa.s.sing through the pupil, could fall upon that part of it. In the middle of the optic nerve is found the branch of an artery, from the internal carotid, which is diffused and ramified in a beautiful manner along the retina. From this artery, a small branch goes through the middle of the vitreous humour, and giving off branches on every side, expands itself upon the capsule of the crystalline lens.

We shall now consider the humours of the eye, which are three in number, the aqueous, the crystalline, and the vitreous; all transparent, and in general colourless; but of different densities.

The aqueous humour, so called from its resemblance to water, fills up all the s.p.a.ce between the cornea and the crystalline humour. It is partly before and partly behind the uvea, and is divided by that membrane into two parts, which are called the chambers of the aqueous humour; which chambers communicate with each other by means of the pupil.

The next humour is the crystalline; it is situated between the aqueous and vitreous humours, and is connected to the choroid coat by the ciliary ligament: it is not the least of all the humours, as has been generally supposed, the aqueous and it being of equal weights; but its substance is more firm and solid than that of the other humours: its figure is that of a double convex lens; but the fore part next the pupil is not so convex as its other side, which is contiguous to the vitreous humour; the diameter of the sphere, of which its anterior segment is a part, being in general about seven or eight lines, whereas the diameter of the sphere, of which its posterior segment forms a portion, is commonly only about five or six lines. It is covered with a fine transparent capsule, which is called arachnoides. This humour is situated exactly behind the pupil, but not in the centre of the eye, as was supposed by Vesalius, being a good deal nearer its forepart. The convexity of its posterior surface is received into an equal concavity of the vitreous humour. It is not of an equal density throughout, but is much more hard and dense towards its centre than externally, the reason of which will appear hereafter. Till we arrive at about our thirtieth year, this humour continues perfectly transparent, and colourless; about that time it generally has a little tinge of yellow, and this colour increases with age.

The third humour of the eye, is the vitreous; it is the largest of all the humours, filling up the whole of that part of the eye which lies behind the crystalline humour. It is thicker than the aqueous, but thinner than the crystalline humour; on its back part is spread the retina, and in the middle of its fore part is a small cavity, in which the whole posterior surface of the crystalline lens lies; this humour is also enclosed in a very fine capsule, called tunica vitrea; this capsule at the edge of the crystalline humour is divided into two membranes, of which the one is continued over the whole anterior surface of the vitreous humour, and lines that cavity into which the back part of the crystalline is received; the other pa.s.ses over the crystalline humour, and covers all its fore part, by which means these two humours are closely connected together. The weights of the aqueous, crystalline, and vitreous humours in a human eye, are, according to the accurate Pet.i.t, at a medium, to each other, as 1, 1, and 25.

It was thought necessary to premise this general description of the structure of the eye, in order that what we are going to add in the remaining part of this Lecture may be the more easily comprehended. A more distinct idea will perhaps be had from a contemplation of the following figure, which represents the section of an eye by a vertical plane pa.s.sing through its centre.

[FIGURE]

EXPLANATION.

NOO represents the optic nerve.

The outmost line ALLB represents the sclerotic coat, and the part ACB the transparent cornea.

The line ALLB, immediately within the former, represents the choroides; the part APB is the iris or uvea, in which the hole at P is the pupil.

The line FOOG is the retina.

The cavity ACBEMDA is the aqueous humour.

DE is the crystalline lens or humour.

The s.p.a.ce DFOOGE, lying behind the crystalline, represents the vitreous humour.

BE and AD is the ligamentum ciliare.

_Nature and Properties of Light._

After this short description of the human eye, I shall next proceed to take notice of some of the properties of light; but shall confine myself to such as are absolutely necessary for explaining the phenomena of vision, as far as that can be done from optical principles.

1. It is, I believe, generally at present agreed, that light consists of exceedingly small particles of matter, projected with great velocity in all directions from the luminous or radiant body. This hypothesis, to which no solid objection has yet been made, appears to be more simple than any other; and is so consistent with all the phenomena yet observed, that we have great reason to think it true: however, as it is not absolutely and directly demonstrated, it may have been wrong in optical writers to have given this hypothesis (for it can only be called a hypothesis) as a definition of light.

2. The s.p.a.ce through which light pa.s.ses is, by opticians, called a medium, and it is observed, that, when light pa.s.ses through a medium, either absolutely void, or containing matter of an uniform density, and of the same kind, it always proceeds in straight lines.

3. Those rays of light which come directly from a luminous body to the eye, only give us a perception of light; but when they fall upon other bodies, and are from them reflected to the eye, they give us an idea or perception of those bodies.

4. When a ray of light pa.s.ses out of one medium into another of different density, it is bent out of its course, and is said to be refracted. We must, however, except those rays which fall in a direction perpendicular to the surface of the refracting medium; as the refractive force acts in the same direction in which those rays move, they will not be turned out of their course, but proceed in the same direction they had before they entered the refracting medium.

When a ray pa.s.ses out of a rarer into a denser medium, it will be refracted, or bent towards a line which is perpendicular to the surface which separates the media at the point where it falls; but when it pa.s.ses out of a denser into a rarer medium, it will be bent from the perpendicular.

5. Whenever the rays, which come from all the points of any object, meet again in so many points, after they have been made to converge by refraction, there they will form the picture of the object, distinct, and of the same colours, but inverted. This is beautifully demonstrated by a common optical instrument, the camera obscura. If a double convex lens, be placed in the hole of a window shutter in a dark room, and a sheet of white paper be placed at a certain distance behind the lens; a beautiful, but inverted picture of the external objects will be formed: but if the paper be held nearer, or more remote than this distance, so that the rays from each point shall not meet at the paper, but betwixt it and the lens, or beyond the paper, the picture will be indistinct and confused.

_Of the Manner in which Vision is performed._

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