Part 3 (1/2)

_Remarks._--_Pteralopex atrata atrata_ is known from four specimens from Guadalca.n.a.l and one from Santa Ysabel (Sanborn, 1931:21).

Sanborn (_loc. cit._) reported that a specimen wounded at night, while feeding on young green coconuts, was the only fruit bat that attempted to attack the collectors. Troughton (1936:348) has suggested, on the basis of his experiences with _Pteropus_, that this behavior probably was a reaction from fear rather than an indication of general aggressiveness on the part of _Pteralopex_.

=Pteralopex atrata anceps= Andersen

1909. _Pteralopex anceps_ Andersen, Ann. Mag. Nat. Hist., ser. 8, 3:266, March, type from Bougainville; 1912, Andersen, Catalogue of the Chiroptera ... British Museum, 1:437; 1936, Troughton, Rec. Australian Mus., 14:348, April 7; 1953, Pohle, Z. Saugetierk., 17:129, October 27.

1954. _Pteralopex atrata anceps_, Laurie and Hill, List of land mammals of New Guinea, Celebes and adjacent islands, p. 40, June 30.

_Specimens examined_ (three males, two females; one skull-only and one in alcohol).--Choiseul in March, 23682; Bougainville in July, USNM 276973-74, USNM 276928, USNM 277112.

_Measurements._--Measurements of three males and one female are, respectively, as follows: Length of head and body, 280, 271, 261, 255; hind foot, 50, 54, 52, 59; ear, 23, 23, 26, 22; length of forearm, 160, 162, 166, 171; greatest length of skull, 77.6, 77.9, 78.9, 77.0; condylobasal length, 74.3, 74.3, 75.5, 73.8; zygomatic breadth, 42.2, 45.4, 43.1, 42.6; breadth across upper canines, 18.7, 21.1, 19.0, 19.0; breadth across first upper molars, 22.2, 25.3, 22.9, 22.0; length of maxillary tooth-row, 29.3, 29.8, 28.9, 28.2; length of mandibular tooth-row, 32.8, 32.8, 32.1, 31.4.

_Remarks._--Heretofore, _Pteralopex atrata anceps_ was not known from Choiseul. The specimen from that island agrees well with specimens in the U. S. National Museum from Cape Torokina, Bougainville.

The type specimen of this subspecies is a subadult and is smaller than the specimens examined by me; Andersen (1912:440) gave length of forearm of the type as 137 (as opposed to 164 in adults). He (1912:438) figured the dent.i.tion of _anceps_ and described the ways in which it differed from the dent.i.tion of _atrata_. Although he (1912:437) concluded that _anceps_ and _atrata_ represented ”two stages of specialization of ...

dent.i.tion,” there apparently are few, if any, dental differences between the two subspecies. Teeth of adults of _anceps_ differ from teeth of the immature type of _anceps_ as follows: in adults the anterior basal ledge of P4 extends onto the l.a.b.i.al surface, whereas in the type it does not; and maxillary and mandibular teeth in adults are s.p.a.ced as in the subspecies _atrata_ (see Andersen, 1912:438, fig. 22) and not crowded as in the type of _anceps_. Distance between individual cheek-teeth apparently increases with growth of the cranium and mandible.

Adults of _P. a. anceps_ that I examined are darker than the subadult type. The mantle in these adults is black, whereas it is seal-brown in the type (Andersen, 1912:439).

An adult female was lactating when obtained on Bougainville in July (USNM 276928).

Key to _Pteropus_ in the Solomon Islands

1. Premolars having distinct basal ledges; molars 2.5-4.0 wide 2

1'. Premolars lacking definite basal ledges; molars 1.0-2.4 wide 14

2(1). Rostrum unshortened (orbit to anterior tip of nasals about one-third greatest length of skull); dorsal surface of tibiae nearly naked 3

2'. Rostrum shortened (orbit to anterior tip of nasals less than one-third greatest length of skull); dorsal surface of tibiae usually at least partially furred 8

3(2). Forearm more than 128 4

3'. Forearm less than 128 5

4(3). Forearm about 155; venter and dorsum nearly black, mantle pale yellow =P. tonga.n.u.s geddiei=, p. 798

4'. Forearm 128-136; venter and dorsum near Mars Brown, mantle Ochraceous or Cream-Buff =P. hypomela.n.u.s luteus=, p. 796

5(3'). Mantle dark, russet or cinnamon, not strongly contrasting with color of back =P. admiralitatum solomonis=, p. 796

5'. Mantle pale, Ochraceous-Buff or Cream-Buff, strongly contrasting with color of back 6

6(5'). Mantle Ochraceous-Orange to Ochraceous-Buff, hairs pale basally; forearm 108-111 =P. admiralitatum goweri=, p. 797

6'. Mantle Ochraceous to Cream-Buff, but hairs dark brown basally; forearm 110-122 7

7(6'). Length of forearm 110-112 =P. admiralitatum colonus=, p. 796